Line 90, 160, 202, 221, 237, 263, 284, 329, 372, 385, 574, 598, 614, 628, 637, 651, 676: The paragraph should be started by special indentation in the first line. Line 55: No abbreviations are allowed in the abstract. Abbreviations should be given explanations when first appearing. Please check and revise, for example: QTLs, qFe1.1, qFe3.1, qFe5.1, qFe7.1, qFe8.1, OsZIP8B, OsZIP6A, qFe3.3, qFe7.3 qZn6, qZn7, qZn2.2, gRMm9-1, qZn8.3 and qZn12.3. Line 64, 65 and 66: add (.) after identified, co-localized and QTLs. Line 101- 102, 96, 99, 103-104, 108, 110, 115, 126, 127-128, 128-130, 130-131, 136-137, 144-145, 146-147, 148, 149-151, 391, 409, 415, 441, 443-444, 460, 501, 529, 543, 632-633, 654, 656, 663, 664, 665-666, 669: (Qui et al. 1995; Ahmed et al.1998; Graham et al. 1999; Gregorio et al. 2000; Anuradha et al. 2012), (FAO 1998), (Allen et al. 2006), (Swamy et al. 2016), (Zhang et al.2009), (Chandel et al. 2011), (Mahender et al. 2016), (Carnahan et al. 1972), (Coffman et al. 1987; Yu et al. 2009), (Stangoulis et al. 2007; Yongmei et al. 2007; Garcia-Oliveira et al. 2008), (Zhong et al. 2011; Huang et al. 2012; Du et al. 2013; Gande et al. 2014; Berhanu et al. 2013), (Lee et al. 2014; Zhang et al. 2014), (Flint-Garcia et al. 2003; Roy et al. 2006; Pradhan et al. 2016), (Zhao et al. 2011; Huang et al. 2015), (Agrama et al. 2007; Huang et al. 2010; Zhao et al. 2011), (Pan et al. 2015), (Zhao et al. 2013), (Kumar et al. 2015), (Muthukumar et al. 2015; Swamy et al. 2017), (Anandanan et al. 2016), (Sellappan et al. 2009), (Jagadeesh et al. 2013), (Cakmak et al. 2004; Morgounov et al. 2007; Johnson et al. 2011), (Ishimaru et al. 2011), (Shabbir et al. 2008; Nagesh et al. 2012; Anjum et al. 2007), (Ravindra Babu et al. 2013; Roy and Sharma 2014; Vishnu et al. 2014), (N’Goran et al. 2000), (Xu et al. 2015), (Brait and Lobreaux 1997; Pflieger 529 et al. 2001), (Kinoshita et al. 2017), (Miura et al. 2010), (Murray and Thompson 1980), (Liu et al. 2005), (Nei 1972), (Evano et al. 2005), (Earl and Von 2012), (Peakall et al. 2012) and (Bradbury et al. 2007) are missing from references.

Line 930-932: add (.) after (2005), Oryza sativa is italic, the correct of (527–553) is (527–534).
Line 713-714: add (.) after (2011), the correct title is (Genetic diversity for iron and zinc contents in a collection of 220 rice (Oryza sativa L.) genotypes), add space after (:). Line 870: add space before (Kiranmayi S), add (L) after (S) in (Kiranmayi SL), add (.) after (2013), replace (e) small by (E) capital letter in (enrichment) and add space between (of) and (iron). Line 867-869: the correct is (Rao DS, Babu PM, Swarnalatha P, Kota S, Bhadana VP, Varaprasad GS, Surekha K, Neeraja CN and Babu VR). the correct title is (Assessment of grain zinc and iron variability in rice germplasm using energy dispersive x-ray fluorescence spectrophotometer (ED - XRF) and replace (46) by (45). Line 844-846: add space after each (,), remove (et) from (Patil et R), add (.) after 2015 then space and add space after (:). Line 858-860: add (.) after 2019, add (294: 963-983) before (https://doi.org/10.1007/s00438-019-01556-w) and add (.) in the end. Line 708: add (Oryza sativa L.) after rice. Line 803-804: add space after each (,), replace ca by Ca, and add space after (.). Line 836-838: the correct arrangement of authors is Nagesh P, Usharani G , Babu VR, Reddy TD, the correct title is (Molecular mapping of high iron and zinc rich regions in rice (Oryza sativa L.) grains using microsatellite markers), the correct is 2: 231-237. Line 890-892: the correct of Anil KB is Batchu AK, add space after (,), add space before QTL, add space before QTL, the correct of concentration is concentrations and O. Nivara is italic. Line 130: remove (;) before Swamy. Line 848-849: the correct is QTLs and indica is italic. Line 137: remove double space before these. Line 927-929: remove (b) from 2012b and the correct is PLOS ONE. Line 149: add ) after 2013. Line 197: add () to Fig. 1. Line 224: add () to Table 2. Line 232: add () to Ho. Line 237: replace stracture by structure. Line 241: add space before 3B. Line 291: add space between axis and 2. Line 294: 1st and 2nd are superscript.
Line 379, 562: replace Figure by Fig. Line 389: add space before ppm. Line 395: add () to Table. Line 437: add space before 2013. Line 518: add () to 2015. Line 620: add () to 2014. Line 919: add Y to Cock JAY. Line 697: replace Agrawal by Agarwal and add SK after Mangrauthia. Line 792-793: add space before Jain, add space after (2014), add space before population, add space after between. Line 997: Fold not bold font.

No Conflicts of interest.